Memecultures have definitely moved onto focus on communities of memers (e.g. private chat servers; rings of Instagram memepages with admins in one chat; podcasters having each other on for special episodes).
Facebook memepages had an unusual alternative to both these silos and the traditional broadcasting model (where one content creator gets seen by a large audience) because it was so easy for them to reach new people in unexpected ways, but that has since been patched away.